University of Illinois at Urbana-Champaign
College of Agriculture
Cooperative Extension Service
Circular 1105
Prepared by A. J. Turgeon, Extension Turfgrass Specialist, and F.
A. Giles, Extension Specialist in Home Landscape.
THE PROPER SELECTION AND CARE OF TURFGRASSES in Illinois depend upon
a knowledge of the environmental adaptation, cultural requirements, and
identification features of a number of grass species. Frequently, questions
arise as to why a particular turf either failed to establish or deteriorated
under certain environmental conditions. Sometimes the answer is that a
poorly adapted turfgrass was planted or that the cultural requirements
of a particular grass were not met. The first prerequisite to determining
the cause of turfgrass problems is an ability to correctly identify different
turfgrass species. The thinning and discoloration of rough bluegrass or
red fescue on exposed sites in summer, the death of annual bluegrass under
conditions of severe drouth and heat stress, the failure of Kentucky bluegrass
to persist in the shade, and the tan color of zoysiagrass in spring are
all situations requiring an accurate identification of the particular grass
present in order to determine the cause of the "diseased" condition of
the turf.
This circular is intended as a guide to learning the important vegetative
features for identifying various turfgrass species found in Illinois. Refer
to circular 1082, "Illinois Lawn Care and Establishment," circular 1076,
"Turfgrass Pest Control," and other extension publications for more specific
information on the culture of turfgrasses.
TURFGRASS MORPHOLOGY
The mature turfgrass plant is made up of fibrous roots and of shoots composed
of elongated and unelongated stems with long, narrow leaves arising from
the stem nodes and terminals. Growth habits (Fig. 1) include bunch-type,
rhizomatous, and stoloniferous. The bunch-type growth habit is generally
found in annual bluegrass, colonial bentgrass, Chewings fescue, tall fescue,
and the ryegrasses. Lateral development of grass plants is by tillering,
a process by which new shoots arise from vegetative buds in the axils of
the leaf sheaths of older shoots. As the enclosing leaf sheath decays,the
newly developed shoot (tiller) is revealed. Bunch-type turfgrasses generally
have poor recuperative ability after injury; reseed bare spots in these
turfs to enhance recovery.
Grasses with a rhizomatous growth habit include Kentucky bluegrass,
Canada bluegrass, and creeping red fescue. In addition to tillering, these
grasses can spread laterally with the growth of below-ground stems called
rhizomes. These develop from the same vegetative buds as do tillers, but
the new rhizome breaks through the enclosing leaf sheath and grows laterally
through the soil. This type of growth habit facilitates rapid growth into
bare areas that have resulted from diseases, insects, traffic, and other
factors.
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Figure 1. Turfgrass growth habits: (a) bun type (insert shows
tillering), (b) rhizornatous, and (c) stoloniferous.
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A stoloniferous growth habit results from the development of stolons.
Like rhizomes, stolons are horizontally growing stems that break through
an enclosing leaf sheath, but their lateral growth is along the surf-ace
of the ground. Stoloniferous grasses include rough bluegrass and creeping
bentgrass. When seeded with other grasses, stoloniferous grasses tend to
become isolated into distinct patches that disrupt the uniformity of the
turf; hence, they are best used in pure stands for specific types of uses
and environmental conditions.
Grasses that have both stolons and rhizomes include the two warmseason
species, bermudagrass and zoysiagrass. As with the strictly stoloniferous
grasses, these are best used in pure stands.
LEAF STRUCTURES
The turfgrass leaf is composed of an upper flattened portion called the
blade
and a lower tubular portion called the sheath. At the junction of the leaf
blade and sheath are the ligule, the collar, and, in some species, auricles.
These are the three principal structural features used in identifying different
turf grass species.
The ligule (Fig. 2) is a tongue-like outgrowth that may be a
fringe of hairs, as in bermudagrass and zoysiagrass, or a membranous structure
of a particular size and shape. Kentucky bluegrass has a very short, flat,
membranous ligule, while creeping bentgrass has a long, pointed, membranous
ligule. A membranous ligule may be smooth across the top, as in redtop,
or jagged, as in colonial bentgrass.
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Figure 2. Ligule structures: (a) fringe of hairs, (b) smooth
membrane, and (c) jagged membrane.
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Figure 3. Continuous collar (left)
and divided collar (right).
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The collar (Fig. 3) is a light green band at the base of the
leaf blade; it varies in size and shape according to species. Kentucky
bluegrass has a narrow, continuous collar, while tall fescue has a broad,
divided collar.
The auricles (Fig. 4) are appendages projecting from each side
of the collar. They may be long and slender, as in annual ryegrass, short
and blunt, as in tall fescue, or absent, as in Kentucky bluegrass.
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Figure 4. Turfgrass leaves with (a) long and slender auricles,
(b) short and blunt auricles, and (c) no auricles.
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Figure 5. Folded (left)
and rolled (right) vernation.
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An additional feature helpful in identifying turfgrass species is thevernation
(Fig. 5), the arrangement of leaves enclosed within the older leaf sheaths.
This characteristic is easily demonstrated with the ryegrasses. Perennial
ryegrass has a folded vernation, with leaf margins meeting but not overlapping,
while annual ryegrass has a rolled vernation, with leaf margins overlapping.
The leaf blades differ considerably among turfgrasses. The bluegrasses
have smooth leaf blades (Fig. 6a), while ryegrasses, fescues, and bentgrasses
have prominent veins on the upper leaf surface (Fig. 6b).
Under moisture stress, the leaf blades of Kentucky bluegrass (Fig. 6c)
and red fescue (Fig. 6d) may fold in response to changes in turgor pressure
of the bulliform cells (large, thin-walled epidermal cells located on the
upper leaf surface). The two translucent lines observed when a Kentucky
bluegrass leaf blade is held up to the light are due to the parallel groups
of bulliform cells located on either side of the midrib.
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Figure 6. Smooth leaf blades of bluegrass (a); ryegrass leaf
blades with prominent veins (b); response of leaf blades to moisture stress
in Kentucky bluegrass (c) and red fescue (d).
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Figure 7. "Boat-shaped" leaf tip of bluegrass.
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The shape of the leaf tip is an important feature for distinguishing
the bluegrasses from other turfgrass species. Bluegrasses have a characteristic
"boat-shaped" leaf tip (Fig. 7), while the leaf tips of most other turfgrasses
are flat and pointed. The shape of the leaf blade moving toward the tip
also helps distinguish Kentucky bluegrass and annual bluegrass, which are
parallel sided, from Canada bluegrass and rough bluegrass, which taper
toward the leaf tip.
BLUEGRASSES (Poa sp.)
Bluegrasses include several species that are important as turfgrasses.
These species differ substantially in general appearance, growth habit,
environmental adaptation, and cultural requirements; however, they all
have certain structural features in common:
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folded vernation (young leaf blades are folded as they emerge from the
enclosing sheaths of older leaves)
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boat-shaped leaf tip (the leaf blade gathers toward the tip to form a three-dimensional
shape like the prow of a boat)
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smooth leaf blades with parallel translucent lines on either side of the
central midrib
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absence of auricles
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membranous ligules
KENTUCKY BLUEGRASS (Poa pratensis L.)
Kentucky bluegrass is the principal turfgrass used in Illinois. It is adapted
to a wide range of environmental conditions and provides an attractive
turf when maintained under proper cultural conditions. It performs best
in open, sunny areas under moderate to high fertilization, moderate mowing
heights 1-1/2" to 2"), and supplemental irrigation during drouth periods.
ANNUAL BLUEGRASS (Poa annua L.)
When maintained under moderately close mowing and frequent irrigation,
annual bluegrass forms a very dense, fine-textured, and vigorous turf during
the cooler months of the growing season. It survives reasonably well on
compacted soils and is well adapted to moist, shaded conditions. However,
its poor tolerance of heat and drouth stress during the summer make it
an undesirable weed throughout most of Illinois. Although rarely planted
intentionally, it is frequently the major component of such intensively
cultured turfs as golf course greens, tees, and fairways.
CANADA BLUEGRASS (Poa compressa L.)
Canada bluegrass forms an open turf of low density and generally poor quality.
It is adapted to cool climates and acid, drouthy soils of low fertility.
It grows best when clipped at 3 to 4 inches.
ROUGH BLUEGRASS (Poa trivialis L.)
Rough bluegrass forms a dense, semi-prostrate turf that is adapted to cool,
moist, shaded environments. It frequently invades other turfs that are
growing on poorly drained, fertile soils during the cooler months of the
growing season. During the summer months, however, rough bluegrass is usually
thinned and discolored because of its poor heat, drouth, and disease tolerance.
As it is not compatible with other turfgrasses, its use is usually limited
to wet, shaded areas where other turfgrass species do not persist.
BENTGRASSES (Agrostis sp.)
The bentgrasses include four species that are important as turfgrasses
in Illinois. Although very limited in their general adaptation and use,
bentgrasses provide the dense, closely clipped turfs important for such
popular sports as golf. Bentgrasses have these structural features in common:
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rolled vernation
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membranous ligules
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flat, tapering leaf blades with pointed tips
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prominent veins on the upper sides of leaf blades
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absence of auricles
CREEPING BENTGRASS (Agrostis palustris Huds.)
Creeping bentgrass forms a smooth, dense turf that, when maintained properly,
is suitable for use in golf course putting greens and tees, bowling greens,
grass tennis courts, and exotic lawns. Its cultural requirements include
close (1/4" or less), frequent mowing; regular applications of fungicides
for disease control; frequent irrigation; and some cultivation or topdressing.
It is not compatible with other turfgrasses and is frequently found as
a serious weed in Kentucky bluegrass lawns.
COLONIAL BENTGRASS (Agrostis tenuis Sibth.)
Colonial bentgrass is not as well adapted to Illinois as to the milder
climates of New England and the Pacific Northwest; however, it is sometimes
found in fairway turfs of northern Illinois golf courses. It does best
at slightly higher (1/2" to 3/4") mowing heights than for creeping bentgrass;
otherwise, their cultural requirements are similar. Once considered an
important component of lawn seed mixtures, the use of colonial bentgrass
is now discouraged because of its tendency to become the dominant species
in a turf.
VELVET BENTGRASS (Agrostis canina L.)
Velvet bentgrass forms a very fine-textured, extremely dense turf when
maintained under close mowing and meticulous care. Its high thatching tendency
and slow shoot growth limit it primarily to the New England States, where
it is especially well adapted. Velvet bentgrass is occasionally observed
as small, tight patches in some shaded putting greens on Illinois golf
courses.
REDTOP (Agrosits alba L.)
Redtop forms an open, coarse-textured turf that is adapted to cool, moist
(or wet) environments. When planted as a component of seed mixtures, it
may persist as unsightly, weedy tufts in the lawn. It is intolerant of
temperature extremes, shade, traffic, or close mowing. Redtop is used primarily
for low-quality turfs for soil stabilization on roadsides, ditches, and
waterways.
RYEGRASSES (Lolium sp.)
Ryegrasses have been used traditionally as nurse grasses in lawn seed mixtures
or for the establishment of temporary lawns. Their quick germination and
vigorous seedling growth make them suitable for use where rapid establishment
and soil stabilization are desired. Because of their intolerance of temperature
extremes, drouth, and certain diseases, ryegrasses sometimes behave like
annuals or short-lived perennials. Ryegrasses have the following structural
features in common:
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membranous ligules
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presences of auricles
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flat, tapering leaf blades with prominent veins on the upper sides, smooth
and glossy on the lower side
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bunch-type growth habit
PERENNIAL RYEGRASS (Lolium perenne L.)
Perennial ryegrass is adapted to cool, moist conditions and may persist
during the winter and summer seasons, provided severe temperatures do not
occur. Because of its rapid germination and vigorous seedling growth, it
is frequently used in seed mixtures as a nurse grass for quick cover. A
fairly wear-tolerant grass, perennial ryegrass is sometimes used for overseeding
athletic field turfs that have been injured from play. The mowing quality
of perennial ryegrasses has generally been poor because of the tough vascular
bundles in the leaves; improved varieties, however, such as Manhattan,
Pennfine, and NK-200 have better mowing quality and provide turfs that
are finer-textured, denser, and darker green than common perennial ryegrass.
ANNUAL RYEGRASS (Lolium multiflorum Lam.)
Annual ryegrass is similar to perennial ryegrass but is less tolerant of
cold, heat, and drouth stresses and typically behaves like an annual or
short lived perennial. Its use as a nurse grass is discouraged because
of its objectionably coarse texture and its severe competition with more
desirable turfgrasses during establishment. It is occasionally recommended
as a temporary lawn grass for soil stabilization until there is a more
favorable season to establish Kentucky bluegrass.
FESCUES (Festuca sp.)
The fescues make up a highly variable genus that includes both coarse-
and fine-textured species. These species differ substantially in growth
habit, environmental adaptation, cultural requirements, and use. As a group
the fescues are adapted to drouthy, infertile soils and possess generally
good wear tolerance. Fescues have the following structural features in
common:
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short, flat, membranous ligule
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prominent veins on upper side of leaf blade
RED FESCUE (Festuca rubra L.)
Red fescue forms a very fine-textured, dense, medium to dark-green turf.
It is best adapted to cool, dry, shaded environments and a low to moderate
intensity of culture. It is relatively intolerant of high temperatures,
poorly drained soils, and saline soils. It is frequently used as a companion
grass in mixtures with Kentucky bluegrass for moderately shaded sites.
There are two subspecies of red fescue that differ principally in growth
habit: creeping red fescue (subspecies rubra) is a rhizomatous grass, while
Chewings fescue (subspecies commutata) is a bunch grass that spreads by
tillering only.
TALL FESCUE (Festuca arundinacea Schreb.)
Tall fescue is a very coarse-textured grass that is adapted to a wide range
of environmental conditions. It has good heat, drouth, and wear tolerance,
and it persists fairly well in moderately shaded environments. It provides
a satisfactory turf on sites that receive a low intensity of culture; however,
it responds well to increased fertilization and irrigation if it is mowed
at a height of at least 1-1/2 inches. Because tall fescue is not compatible
with other turfgrasses, it should be planted alone at high seeding rates
(6 to 8 pounds per 1,000 square feet) to ensure the best quality turf.
BERMUDAGRASS (Cynodon dactylon (L.) Pers.)
Bermudagrass is a warm-season species used occasionally as a turfgrass
in southern Illinois. It is also found as weedy patches in bluegrass and
bentgrass turfs. It forms a dark-green, dense, vigorous turf of medium-fine
texture. Its heat and drouth tolerance are excellent, but poor cold tolerance
and winter discoloration severely limit the practical use of bermudagrass
in Illinois.
ZOYSIAGRASS (Zoysia japonica Steud.)
Zoysiagrass is a warm-season turfgrass frequently seen in lawns and on
golf course fairways in southern Illinois. It forms a dense, tough, slow-growing
turf and has good heat, drouth, and cold tolerances. Its tan appearance
while dormant during the cool months of the year (October to May) may be
objectionable, but green dyes may be used to provide artificial color.
The invasion by cool-season grasses and broadleaf weeds during winter should
be controlled with herbicides in order to maintain a uniform turf. Zoysiagrass
is propagated by plugs or shredded sod and is very slow to establish, usually
requiring two to three years.
The Illinois Cooperative Extension Service provides equal opportunities
in programs and employment.
Urbana, Illinois
Reprinted July 1987
Issued in furtherance of Cooperative Extension Work, Acts of May 8
and June 30, 1914, in cooperation with the U.S. Department of Agriculture,
WILLIAM R, OSCHWALD, Director, Cooperative Extension Service, University
of Illinois at Urbana-Champaign.
To 5-84-2SM
3M-6-87-67569